MindMap Gallery Orchidaceae Morphology 1
Vegetative organs, orchids are a very special class of plants with many unique morphological characteristics, such as roots, stems, leaves, flowers and fruits.
Edited at 2024-02-11 12:11:35Avatar 3 centers on the Sully family, showcasing the internal rift caused by the sacrifice of their eldest son, and their alliance with other tribes on Pandora against the external conflict of the Ashbringers, who adhere to the philosophy of fire and are allied with humans. It explores the grand themes of family, faith, and survival.
This article discusses the Easter eggs and homages in Zootopia 2 that you may have discovered. The main content includes: character and archetype Easter eggs, cinematic universe crossover Easter eggs, animal ecology and behavior references, symbol and metaphor Easter eggs, social satire and brand allusions, and emotional storylines and sequel foreshadowing.
[Zootopia Character Relationship Chart] The idealistic rabbit police officer Judy and the cynical fox conman Nick form a charmingly contrasting duo, rising from street hustlers to become Zootopia police officers!
Avatar 3 centers on the Sully family, showcasing the internal rift caused by the sacrifice of their eldest son, and their alliance with other tribes on Pandora against the external conflict of the Ashbringers, who adhere to the philosophy of fire and are allied with humans. It explores the grand themes of family, faith, and survival.
This article discusses the Easter eggs and homages in Zootopia 2 that you may have discovered. The main content includes: character and archetype Easter eggs, cinematic universe crossover Easter eggs, animal ecology and behavior references, symbol and metaphor Easter eggs, social satire and brand allusions, and emotional storylines and sequel foreshadowing.
[Zootopia Character Relationship Chart] The idealistic rabbit police officer Judy and the cynical fox conman Nick form a charmingly contrasting duo, rising from street hustlers to become Zootopia police officers!
Orchidaceae Morphology 1
Image Source
01
https://bklynorchids.com/2011/06/03/orchid-of-the-week-platystele-jungermannioides/
01Platystele jungermannioides smallest orchid
The smallest orchid is not yet certain
02
http://orchids.la.coocan.jp/Grammatophyllum/Grammatophyllum speciosum/Grammatophyllum speciosum.htm
02Grammatophyllum speciosum (giant orchid)
03
https://www.rgbstock.es/photo/mz21AQe/Vanilla Vine
03 Vanilla
Overview
largest longest and smallest
Refer to "Orchid Museum"
Overall form
Stamens grow on one side of the flower
No matter the quantity is 1, 2 or 3
Stamens and stigma at least partially connate
A complete union is called a column
Seeds are small and numerous
The original orchid seeds were slightly larger, but still very small compared to other plants
The flower has a specialized lip
Opposite of fertile stamens
Occasionally less specialized, similar to the other two from a round
As the flower develops, it twists into a position with the lip underneath.
The rostellum (the tongue-like protrusion in front of the stigma in the stamen) is involved in the pollination process
Pollen clumps together
It is an important characteristic to adapt to pollination by insects and birds.
morphology
Important ways to distinguish different groups of orchids
Morphological details of orchids are fundamental to discussions of ecological diversity
important for understanding its evolution and pollination
growth habits
figure 1
A. The leaves of the original orchid are arranged in a spiral, pleated fan-shaped leaves. New branches sprout from the base of the previous year's branch buds every year, and the bases of the new branches over the years together form a rhizome.
Whole plant model
A is the basic pattern of the primitive orchid family, without pseudobulbs and terminal racemes.
The direction of morphological evolution includes several points
Inflorescence position changes
Top student
A,F-K
Axillary
the remaining
shortening of stem
shortened internode
O Phalaenopsis
The internode is extremely shortened to the tip
CEKL pseudobulb
Internodes are extremely shortened to the root neck
FG Paphiopedilum-shaped
thickening of stem
Spindle-shaped enlargement
BDIJ dendrobium-shaped
Root position changes
on rhizomes
on the stem
NP
The roots of P can photosynthesize
leaf position
Terminal leaf cluster
J
Terminal simple leaf
E
Jisheng
CFG
With or without distinct independent petioles
A-M are all sympodial branches
N-P is a uniaxial branch
A spiral birth
After that, there are two rows of each other.
B corm is formed by multiple nodes
C is formed from one section
D,I,J pseudobulbs are formed by multiple nodes
E, K, L section is formed
F tuberous roots are enlarged
G root neck position root
Both are earth-born
Uncommon type
A
Pseudobulbs continue to grow after flowering on the pseudobulbs
Scaphyglottis
B
Pseudobulbs growing on bi-row alternate leaf stems
Individual Maxillaria
C
No pseudobulbs, but fleshy leaves
Subtribe Pleurothallidinae
root
No fine, fibrous root system
There is obvious root cover (velamen)
Tissue with thickened cell walls that dies after maturity
The simplest root integument is a layer of cells, similar to the epidermis in all aspects
More commonly composed of 2-18 layers of cells
Root cover is a product adapted to epiphytic life
Some studies have found algae in the root cover, which can achieve the function of nitrogen fixation. The root cover naturally contains mycorrhizal fungi, so the root cover is a quite complex micro-ecological environment.
exodermis
The cell layer between the root integument and cortex
Long, thick-walled cells lacking cytoplasm
and short living channel cells
The root tip mainly appears green when exposed to light
Mature root cortical cells contain chloroplasts
Although the root cover obscures the green color, some Vanda species rely entirely on cortical photosynthesis
Very short stems with only scale leaves
Taeniophyllum
There are regular round holes on the outer wall cells of the root, and there are also patchy cell groups without holes.
growing from stems
Most roots of orchids are fleshy
A few are uneven in thickness, and the thicker parts are called nodular tuberoids.
Used to distinguish from those whose entire root is thick
Cheirostylis (now a synonym of Cheirostylis)
Very few roots, only fleshy root ridges attached to the rhizome
Each ridge is anatomically a root, with root hairs on the outer surface and a root stem on the inner surface.
The result of extreme root shortening
root-stem tuberoid
The sheath of the root structure, the core of the stem structure surrounding the terminal bud
hibernation structure
droper/sinker
During the growing season, new branches form from the top, and lateral buds develop into new roots.
Sometimes it extends from the leaf base and is buried deep in the soil
In Orchideae (Red Door Orchid Family) and Diurideae (Double-tailed Orchid Family) this structure is multi-pillar
This means that there are multiple conjoined roots in one layer of skin.
This structure in the orchid family is ellipsoidal
There are usually two growing seasons, the old ones are being consumed and the new ones are being created
A pair of testicles shaped like a mammal
orchis refers to the name of this thing in Greek
And eventually became the family name of the plant
tuberoid
A-D
Transverse and longitudinal sections of single/multidimensional tube bundles
Morphological subdivision
E
root-like
F
spindle
G
palmate
H
with handle
I
sessile oval
Root systems buried in the substrate or completely suspended are often cylindrical, but those crawling on the surface are mostly flat or flat underneath.
In some taxa, all roots are flat regardless of whether they are attached or not
Epiphytic and lithophytic taxa may have thick fleshy roots and thin stems and leaves
In some taxa, in addition to the normal root system, non-absorptive roots tend to grow upward from the substrate
Possibly due to accumulation of debris from resorbed roots
Root systems usually grow on the roots
But adventitious buds will also grow on the roots
This type of root system looks like an elongated rhizome.
But the structure is very different
stem
Vascular bundles are denser at the periphery of the stem and are distributed in the parenchyma tissue
Linear and slender
lignification
Soft and juicy
Rhizome
compound organ
Made up of the base of the trailing branches
The horizontal part is made of thickened hard wood
Individual lateral buds in several nodes develop to perfection and form new branches
For taxa with lateral inflorescences, the inflorescence axis often grows from this position.
Rhizome also refers to the horizontal stolons of the Goodyerinae subtribe, because in this group there is no clear boundary between the rhizome and the aerial horizontal stolons.
Single-branched orchids have no rhizomes
The term secondary stem is unreasonably used to describe the branches above the rhizome.
Morphologically, only the first branch produced from a seedling can be called a primary stem, all other stems are secondary.
Aerial stems or erect vegetative shoots are more accurate, albeit odd, terms in the case of terrestrial orchids.
In epiphytic orchids, the entire plant is aerial, and the plant branches (vegetative shoots) may be horizontal or climbing (horizontal, pendant)
It is also difficult to give the term secondary roots an alternative meaning because the term was coined by accident rather than planned.
Corms and pseudobulbs
There is no clear boundary between the two
corms refers to underground storage stems
pseudobulbs refers to the thickened stem structure of epiphytic orchids
Not a tuber
Tuber (underground structure)
Not a bulb
Bulbs refer to structures wrapped by onion-like scale leaves (thick leaf bases)
And the normal function of the stem is not affected
pseudobulb
Formation of multiple internodes (heteroblastic ~ heteromorphic pseudobulb) or single internode (homoblastic ~ homomorphic pseudobulb)
The knobby pseudobulbs will have leaves growing on them, or growing on the top.
In some taxa, pseudobulbs are often wrapped in leaf sheaths, with only one scale leaf on the top.
Pseudobulbs are not suitable in uniaxially branched orchids, so uniaxially branched orchids have other storage organs such as stems and leaves
In Bulbophyllum minutissimium and B.odoardii there is a chamber at the top of the pseudobulb with a small opening to the outside
Former photo
The opening of the former is blocked by scale-like leaves or blades
The interior of the chamber is rich in pores for exchanging outside air
The latter leaves also have stomata (stomata)
Pseudobulbs can photosynthesize even after their leaves fall off
Eria bractescens and its relatives have another type of chamber
Inflorescence buds sprout from the base of the columnar hole on the pseudobulb and are usually closed by leaf sheaths, which are closely attached to the surface of the pseudobulb. When blooming, the inflorescence needs to break through the surface of the pseudobulb, sometimes forming a trapdoor. When the old inflorescence rots, it will leave a neat columnar hole on the pseudobulb.
leaf
In many orchids, only scale leaves or leaf sheaths are attached to the rhizome.
The leaves of most orchids are standard monocot leaves
parallel veins
A
But in Epistephium and Clematepistephium they are reticular veins
Epistephium matogrossense
B
Pachyplectron
with sword-shaped leaves
F
Acianthus
Deeply divided palmate leaves
So much so that it looks more like the Ranunculaceae family.
C
Catasetum
fan folded into leaf sheath
D
Stanhopea
Fan-folded to a single petiole
E
Monophyllorchis
Fan-folded, heart-shaped leaves
G
Thelymitra spiralis
tip twist
H
Dendrobium cucumerinum
fleshy and protruding
Many times the stem or pseudobulb only has one leaf
But looking at the origin of the scale leaves or leaf sheaths reveals a bi-row alternate structure.
In primitive orchids, there is spiral phyllotaxis
Shortened internodes make the plant look like more leaves are emanating from the same level
Codonorchis
Codonorchis lessonii
Isotria
Isotria medeoloides
vernation (phyllotaxis within buds) and folding (leaf folding)
A commonly used feature in orchid morphology is the folding or rolling of leaves during development.
The original group is rolled/convolute
AB
Duplicate patterns are more common in taxa represented by epiphytic orchids.
C
Often bifolded when mature (conduplicate)
Fold at the center line and cut vertically into a V shape
Leaf veins are similar in size and not prominent
E
Curled leaves may become fan-folded or bifold as they develop
When there are fan folds or pleats, part of the leaf veins will protrude, and these protruding parts are creases.
D
Bifold leaves evolved from fan-folded leaves in some taxa
The leaves in some fold-leaf taxa are not exactly one type or a certain standard type.
Chondrorhyncha
Chondrorhyncha lendyana
cross section
columnar
G
triangle
HI
varying degrees of terminal flattening
Fan folding leaves are relatively thin
Compared with the double folded leaves that grow in curls
However, bifold leaves that grow overlapping may be thin or extremely fleshy.
In extreme cases triangular and cylindrical cross-sections
Many orchids have flat leaf tips instead of ventral and abaxial HI
Often called equitant
sheaths and petioles
No leaf sheath, insufficient support capacity of intervening meristem
In other cases, the formation of sheaths is independent of the leaves, especially when the sheaths on rhizomes and inflorescences are very small.
The leaf sheath surrounding the pseudobulb in Teuscheria is very hard and maintains the shape of the pseudobulb. The inner pseudobulb can still maintain its shape after shrinking.
Teuscheria wageneri
The leaf base of some orchids will form a long and narrow cylindrical petiole, such as the Stanhopeinae subfamily (Stanhopeinae).
articulation (connection)
The abscission layer is lacking in many terrestrial orchids.
Leaves simply rot in situ
Most members of the Shulan and Vanda subtribes have
disappears again in highly specialized miniature orchids
In most cases, the abscission layer is between the leaf sheath and the blade.
In a few cases, such as Teuscheria and Oecoclades (no Chinese translation, African Spotted Orchid), the abscission layer is at the lower part of the petiole.
At first glance one would think that the structure at the lower part of the node is the tip of the pseudobulb, but some species of this genus have two leaves on one pseudobulb, each with a joint in the middle of the petiole.
stomata and subsidary cells
The morphology of guard cells and accessory cells is an important basis for plant classification
There are currently three modes in Orchidaceae
no accessory cells
A-C
All Orchidinae
Orchidoideae
Some Cyperoideae
Cypripedioideae
some orchid subfamily
Apostasioideae
Julanya
Pogoniinae
Canna
Epistephium
Mesoperigenous type stomata, D
Among the accessory cells in the stomata, there is a stomatal type that co-occurs from the same mother cell as the guard cells.
another from a neighboring cell
This is the only example among monocots
Cylinaceae
Spiranthoideae
Peripheral development (EF) with trapezoid cells
In this pattern, cells on one side of the meristem (each in a different row) divide diagonally to produce ladder-shaped cells.
Ladder cells may develop into accessory satellite cells or divide into 1-2 accessory satellite cells
Nearly all Epidendroideae and Vandoideae, and some Triphoreae
There are also polar accessory cells at the ends of the stomata, which are also differentiated from adjacent cells.
Accessory satellite cells exist in both Cypripedioideae and Apostasioideae, but developmental studies are unclear
Four or more accessory cells are characteristic in more evolved taxa. The pattern of two accessory cells or no accessory cells is found in primitive orchids. Williams (1979) considered the lack of accessory cells to be a primitive trait in Orchidaceae.
inflorescence
Orchidaceae inflorescences are usually racemose, with flowers axillary on the inflorescence axis and blooming from bottom to top.
Several kinds of orchids, such as Orchis simia (a certain red door orchid), have the top flowers blooming first. Although the flowering sequence has been adjusted, they are still racemes.
B
racemes (panicles) of racemose branches
D
Inflorescence usually with only one flower
In all orchids the flowers are opposite to the bracts
Bracts may be inconspicuous or large and colorful
Large and colorful in Cyrtopodium or Lockhartia
Cyrtopodium flavum
Lockhartia lunifera
Flowers spiral on the inflorescence axis
But the bracts and flowers are dichotomous and alternate in some taxa.
In some cases the flowers are whorled
Such as Chamaeangis and Oberinia
Oberonia padangensis
Chamaeangis vesicata
Inflorescences may arise from any part of the stem
The inflorescence in the original group is terminal and is an extension of the central axis of the branch.
In other cases, the inflorescence is lateral or basal
In view of the growth habit of monocots, the inflorescences are always lateral.
Sometimes the inflorescence is extremely shortened, with flowers blooming sequentially on the short axis.
Epidendrum nocturnum
Systeloglossum ecuadorense
Bromheadia petuangensis
Thrixspermum
Sometimes the flowers are produced simultaneously or nearly simultaneously, in tight clusters, as in Elleanthus or Glomera
elleanthus vernicosus
Glomera montana
Sigmatostalix produces each flower not from a single bract, but from a cluster of bracts, and thus may represent a more complex type in extremely shortened inflorescences
Sigmatostalix minax
Uncommon inflorescence types such as
The inflorescence of Lockhartia is either cymose or determineate
Common in dicots, uncommon in monocots
The first flower is terminal
Then the subtending 1 to 2 buds will produce new branches ending in flowers, and the cycle repeats.
The ancestor of this group may have been a single-flowered inflorescence
Evolution favors larger inflorescences
This is achieved in the form of cymes
In the distantly related genus Notylia, flower clusters often arise from the base of older inflorescences. If the number of this flower cluster is reduced to 1, it can be compared with the inflorescence of Lockhartia.
Regular inflorescences arise from the central axis of leaves or bracts
one exception
In Dichaea the single-flowered inflorescence arises directly opposite the leaf base
Dichaea ancoraelabia
Two possible explanations
In many cases the inflorescence opposite the leaf represents the end of a synaptic branch, meaning that each flower is terminal and the internode of the next stem segment grows from the central axis of the opposite leaf.
The other is that the axillary buds come up when the internodes elongate and occupy the opposite position, but in fact they are still axillary.
Inflorescences of Luisia teretifolia are formed in the axils
Wirth, 1964
But it is produced in the middle of the internode
So it seems clear that it is generated on the opposite central axis
Opposite flowers and leaves also exist in other genera and species
Luisia tristis